Upgraded versions of the 4.1 and 5.1 are, in many cases, the fastest Apple computers you can buy today. These are the last highly upgradeable and modifiable desktop computer that Apple sold before moving to the much-maligned “black trash can” design that is sold today and hasn’t changed significantly since 2013.
The Mac Pro 4.1 and 5.1 are known in the community as the “cheese grater” Mac Pro towers.